Wow, great work on that trunk and on those leaves! I’m checking out the file now and it looks *gorgeous*.
We do still need to refrain from using any emission, since keeping shadows consistent throughout all the assets will be super important.
Using some SSS was a great idea. I like it! Turning on Subsurface Translucency in the material helps even more.
I would recommend adding some specular back in, since that helps give it shape.
Mixing in some translucency like this also helps to make it more painterly:

Lastly, even though the vertex count is pretty low thanks to instanced collections, the amount of transparent planes makes the compiling take a bit longer than is probably necessary. See if the middle of each clump can be an opaque sphere of leaves and delete all the planes inside of it to help this render faster:

Keep up the good work!
Edit: Oh, and I do still think that the scale of the leaves is a bit off. I think the size of the clumps you have is perfect, but in terms of the leaves themselves, something like this would work much better:
