Hi Mike,
Never do that (Applying All Transforms) again! (You got lucky here, because your model was not Rotated or Translated.)
The rules for Meshes are:
Always (and this is very important) Apply the Scale.
Don’t Apply the Rotation, unless you absolutely have to.
And never Apply the Location.
You might eventually run into a situation, where you may break one of those rules, but that is pretty rare.
